Am not familiar with Wills' argument, but PBS had a show about the Scopes trial some time ago (from the "American Experience" series, I believe) that was very sympathetic to Bryan. The show's POV was that Bryan wasn't opposed to teaching actual Darwinist theory in schools as much as he was worried about the using that teaching to indoctrinate students in support of Social Darwinism, eugenics, etc. Looking at how well the US has turned out as a "meritocracy" run by high-IQ psychopaths, I think Bryan had a point.
Carl
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